The Eye
Main features
Light control
- The iris controls the size of the pupil to regulate the amount of light entering the eye
- The circular and radial muscles in the iris work antagonistically to adjust the size of the pupil
Focusing
- The lens changes shape to focus light rays and allow clear vision at different distances
- The ciliary muscles attached to the lens control its shape and adjust the degree of refraction
Light receptors - rods and cones
Rods are sensitive to low light conditions and provide black and white vision
Cones are sensitive to colour and function in bright conditions, providing detailed vision
The fovea, located in the center of the retina, contains only cones and provides the sharpest vision
Structure and Function
Aqueous and Vitreous Humour
- Aqueous humour is a watery fluid that maintains the shape of the front part of the eye
- Vitreous humour is a jelly-like fluid that maintains the shape of the back part of the eye and supports the lens
Blind spot
- A small area on the retina where the optic nerve leaves the eye
- No light-sensitive cells are present, so any image falling here is not seen
Ciliary Body
- Contains the ciliary muscle, which is attached to the lens
- The ciliary muscle changes the shape of the lens to allow for focusing at different distances
Cornea
- A transparent tissue that covers the front of the eye
- It refracts (bends) light more than any other part of the eye
Iris
- A muscular layer that contains circular and radial muscles
- It controls the size of the pupil and the amount of light entering the eye
- Also contains a pigment that absorbs light
Lens
- A transparent biconcave structure that refracts light passing through the eye
- It can change shape to adjust the degree of refraction and focus light
Optic Nerve
- A bundle of sensory nerve fibres that transmit visual signals from the eye to the brain
Choroid
A layer of pigmented cells between the sclera and retina
It absorbs light and prevents internal reflection within the eyeball
Contains blood vessels that supply the retina and other layers with blood
Retina
The inner layer of the eyeball that contains rods and cones
Also contains bipolar neurons and ganglion cells that transmit visual signals to the brain
Sclera
- The tough, opaque outer covering of the eyeball
- It protects the eyeball against mechanical damage and provides attachment points for eye muscles
Conjunctiva
- A thin, transparent membrane that protects the cornea against damage
- Tears lubricate the surface of the conjunctiva
Adjustment of Pupil Size
Iris Muscles
The iris consists of circular and radial muscles that control the size of the pupil
The circular muscle constricts the pupil, while the radial muscles dilate the pupil
These muscles work antagonistically to adjust the size of the pupil
Reflex response
The adjustment of pupil size is a reflex response controlled by the autonomic nervous system
The iris responds rapidly to changes in light intensity to protect the sensitive rod and cone cells
Very bright light can overstimulate the cells and cause damage
Accommodation
Focusing Light
The eye adjusts the refraction of light to focus on objects at different distances
Light is refracted/bent as it passes through the cornea and lens
The cornea provides most of the refraction, but the lens can adjust the degree of refraction
Lens Adjustment
The lens changes shape to adjust the degree of refraction
For distant objects, the ciliary body relaxes, causing the lens to become thinner and less converging
For near objects, the ciliary body contracts, causing the lens to become thicker and more converging
Reflex Actions
- Accommodation and the control of light entering the eye are reflex actions
- They are automatic responses and not under voluntary control
Rods and Cones
Structure
Both rods and cones have an outer segment containing a photosensitive pigment
They also have an inner segment with a pair of cilia leading to the cell's nucleus
Rods and cones synapse with bipolar neurons, which then synapse with ganglion cells
Differences
Rods are more numerous and evenly distributed throughout the retina, while cones are concentrated in the fovea
Rods are very sensitive and operate in dim light, while cones operate only in bright light
Rods are insensitive to colour and provide monochromatic vision, while cones are sensitive to red, green, or blue light and provide colour vision
Rods have poor resolution and visual acuity, while cones have good resolution and high acuity
Pigments
Rods contain rhodopsin
Cones contain iodopsin
Rhodopsin is more sensitive than iodopsin and is readily broken down by light
Synapses and visual acuity
Several rods synapse on one bipolar neuron, allowing for retinal convergence
This increases the sensitivity of rods to low light intensity but decreases visual acuity
Cones synapse individually with bipolar neurons, providing high visual acuity and precise colour vision
Trichromatic Theory of Colour Vision
Types of cone cell
There are three types, each sensitive to a different wavelength of light
Blue cones respond to blue light, red cones respond to red light, and green cones respond to green light
The perception of color depends on the relative stimulation of these three types of cone cells
Colour perception
Pure blue, red, or green light will only stimulate the corresponding cone cells
Most light is a combination of different wavelengths, and the degree of stimulation of each type of cone determines colon perception
For example, yellow light stimulates both red and green cones, resulting in the perception of yellow
Structure of the Retina
Layers
Contains millions of light-sensitive cells (rods and cones) and the neurons with which they synapse
Rods and cones form a layer inside the choroid, followed by a layer of bipolar neurons
Beyond the bipolar neurons, there is a layer of ganglion cells
The axons of ganglion cells form the optic nerve, which carries visual signals to the brain
Inverted Arrangement
The retina is said to be inverted because neurons lie over the photoreceptor cells
Light has to pass through the neurons before reaching the rods and cones, which can compromise light detection
The blind spot is where the optic nerve leaves the eye, and no light-sensitive cells are present
Retinal Convergence
Occurs when several rods synapse on one bipolar neuron
This allows for summation of sub-threshold stimuli and increases visual sensitivity
Cones synapse individually with bipolar neurons, providing higher visual acuity
Binocular Vision
Advantages
Binocular vision is the ability to see with both eyes and create a single 3D image
The visual fields of both eyes overlap, allowing for accurate judgment of distance and depth perception
Binocular vision provides larger visual field, better visual acuity, and stereoscopic vision
Predatory Species
Predatory species, like humans and foxes, have eyes positioned at the front of their head for binocular vision
Binocular vision is important for accurately judging distance and position for hunting
Humans have a binocular visual field of 180°
Prey Species
Prey species, like rabbits, have laterally placed eyes on the side of their head
This gives them a wider field of view to detect potential predators
Prey species do not prioritize 3D vision and do not have stereoscopic vision