Features of exchange surface: maintenance of steep diffusion gradient (e.g. mass transport system / rich blood supply by dense capillary network or ventilation by movement of water/air/environmental medium ), short diffusion pathway(e.g. thin/flat organism, branching network of vessels, one-cell thick endothelium),large surface area(e.g. capillary network, rbcs move thru in single file, microvilli, spiracles and branching tracheoles)