Plant And Animal Responses
An adaptive advantage refers to any adaptation that promotes survival of the organism, increasing its chances of breeding successfully and therefore contributing its successful alleles to the gene pool.
To survive, plants and animals need to be able to sense and respond to their external environment.
Abiotic factors - non-living factors
Biotic factors - living factors
Temperature
Light intensity
Salinity
Humidity
pH
Water
Oxygen levels
Carbon dioxide levels
Mineral levels
Substrate
Competition
Mutualism
Exploitatiom
Predation
Parasitism
Animals
Orientation In Space
Taxes
Kineses
Homing
Migration
Plants
Orientation In Space
Tropisms
Nastic Movements
Orientation In Time
The environment displays regular cycles based on the movement of Earth and the moon.
Daily
Lunar
Tidal
Competition For Resources
Interspecific Relationships
Exploitation
Mutualism
Herbivory
Predation
Parasitism
Intraspecific Relationships
Territoriality
Hierarchical behaviours
Co-operative interactions
Reproductive behaviours
The ecological niche of an organism is the way it has adapted in response to the habitat in which it lives.
The habitat
Its adaptations - structural, behavioural, physiological
The fundamental niche is that which the organism would occupy if all the necessary environmental conditions were met.
The realised niche is the actual niche the organism occupies
Limits to the fundamental niche are set by the organism's tolerance to abiotic factors
The limits to the actual niche are typically set by biotic factors, particularly interspecific competition
Tropisms are growth movements either towards (positive, +ve) or away from (negative, -ve) an external stimulus.
The direction of the stimulus determines the direction of growth.
Types of Tropisms
Phototropism (stimulus is light)
Chemotropism (stimulus is a particular chemical such as toxins, acids)
Geotropism/gravitropism (stimulus is gravity)
Thigmotropism (stimulus is touch)
Hydrotropism (stimulus is water)
Experiments into the control of tropisms indicate that plant hormones such as auxin are responsible. Most investigations have revolved around phototropism and geotropism.
Auxins (eg, IAA) are produced in the cells of growing tips of stems and roots.
The diffuse down from the tip and promote elongation of cells (hence the stem grows in length).
Increasing the concentration of auxin up to a certain level increases cell elongation in stems; above this level elongation is inhibited.
If auxin concentrations are unequal on either side of the stem, cells will elongate more on one side of the stem than on the other. This results in stem bending.
In roots, high concentrations of auxin inhibit cell elongation. Auxin concentrates on the lower side of the root, due to gravity, inhibiting cell elongation on this side. This results in the root bending downwards and therefore into the soil, ie, positive geotropism.
Nastic movements are rapidly reversible movements of part of a plant in response to changes in the intensity of a stimulus (not its direction).
Types Of Nastic Movements
Thigmonasty (stimulus is touch)
Thermonasty (stimulus is temperature)
Investigations indicate that rapid movement/collapse of parts of the plant may be the result of changes in turgor pressure in the cells at the base of the stem of the leaf.
Taxes are (rapid) movements either towards (positive, +ve) or away from (negative, -ve) an external stimulus.
The direction of the stimulus determines the direction of movement.
Types Of Taxes
Chemotaxis
Geotaxis
Thermotaxis
Phototaxis
Hydrotaxis
To orientate towards or away from the stimulus, the organism either determines the direction of the stimulus by moving a single sense organ and comparing relative intensities (klinotaxis), or the organism determines the direction of the stimulus by comparing using two or more sense organs or either side of its body and comparing the intensity of the stimulus (tropotaxis).
Kineses are non-directional movements in response to a stimulus, with the intensity of the stimulus determining the rate of movement (the direction of the movement in random, bearing no relation to the direction of the stimulus).
Orthokinesis, in which the intensity of the stimulus determines the speed or rate of movement - the rate is faster in unfavourable conditions and slower in favourable conditions
Klinokinesis, in which the intensity of the stimulus determines the rate of turning when moving - the rate is faster in unfavourable conditions and slower in favourable conditions.
The adaptive advantage of tropisms, nastic movements, taxes and kineses is to move the organism into favourable conditions (and/or out of unfavourable conditions/harmful conditions)
Homing is a response in which an animal regularly returns to a nest site or a breeding nest/colony.
To do this successfully, the animal must be able to navigate and requires an internal clock,.
Migration is a response in which members of a species move from one geographic location to another isolated geographic location.
Typically, migration is an annual event in which animals migrate between breeding and feeding grounds. This may be between two different parts of a country or between two different parts of the world.
For some species, return migration occurs at different stages of their life cycle and not annually.
Migration requires the ability to navigate, as well as the presence of an internal clock.
Migration is innate (adults do not teach their young where to go or how to get there). However, migratory experience over the years can improve success rate.
Animals need to prepare for migration
Laying down of fat layers to ensure energy supplies for the journey.
Μoulting of feathers and replacement with new ones (especially flight feather) to ensure maximum flight efficiency. The new feathers may include breeding plumage for the spring migration.
Migration involves risks to individuals.
Insufficient fat deposits to make the lengthy journey
Feathers in poor condition so the journey cannot be completed
Predation during the journey (predators will congregate at sites that the birds will fly over during migration periods)
Birds too weak/ not healthy enough to complete the journey
Storms blow the birds off course / kill them
Migration provides distinct benefits
Avoiding extremes of winter climate at each location (with reduced / no available food)
Plentiful hours for feeding at each location
Promoting survival / breeding success
Young grow rapidly and are strong
Abundant food supplies reduce infraspecific competition, increasing survival
For long-distance migration to have evolved and continue to exist, the advantages/benefits must outweigh the disadvantages/costs. Migration offers greater reproductive success compared with non-migration.
Navigation may involve several different methods.
Landmarks - the animal recognises familiar landmarks and uses these to guide it to its destination. This is more common when the animal is close to home.
Solar navigation (sun compass) - the sun moves across the sky from east to west during the day and any animals are able to use this to navigate. As the sun is so far away, if the animal is able to keep a set angle to the sun when moving, it will move in a straight line.
Since the sun moves across the sky during the day, a long-distance migrant must compensate for the sun's movement. Therefore an internal clock is essential to keep track of time and allow the animal to change its orientation to the sun and therefore keep on course to its destination.
Stellar navigation (star patterns) - animals can use star patterns such as the constellations when moving at night. An internal clock is also required, to compensate for movement of the stars.
Magnetic fields - many animals are known to use Earth's magnetic fields to successfully navigate.
Chemical trails - animals may use scent trails to find their way to a destination
Sonar (sound reflection) - the ability to navigate using echolocation, emitting sound waves that bounce off objects.
Typically, a combination of methods may be used by an animal to successfully find its destination.
Can give plants support. Saves energy and materials by not needing to produce large amounts of the fibrous cells/ tissues normally needed to support the stem. This energy can then be used for other life processes.
Maximises higher light intensity to maximise photosynthesis and therefore increase growth.
Being higher gives greater exposure to pollinators, increasing opportunities for successful pollination.
Increased height about the ground also provides better dispersal opportunities for seeds.
Photokinesis is a response to changes in light intensity, with the rate of movement determined by the light intensity.
Higher light intensity = higher temperature = higher chance of dehydration
Migration cues
Change in photoperiod. The reason that the cue is change in photoperiod is that this is consistent from place to place and throughout the year.
Other cues that might be used are variable and do not produce a consistent trend, therefore migration may occur at not the most opportune time of the year.
Breeding in areas with longer photoperiod
Abundant food
Weak individuals that do not complete the migration have their alleles removed from the gene pool; only strong/best-adapted individuals get to compete the route and successfully breed.
Cells lose water in osmosis and therefore lose rigidity
Evolved as a response to reduce herbivory.
Revert to the horizontal to gain full exposure to light, allowing maximum photosynthesis
Earth rotates on its axis ever 24 hours
Annual
Resulting in regular seasonal cycles with changes in photoperiod and temperature
The gravitational pull of the moon orbiting Earth results in tides (with approximately 12 hours between successive high tides and low tides).
Organisms use environmental cues to set their activity patterns/rhythms.
Animals that are night active are nocturnal
Animals that are day active are diurnal
Animals that are active at dawn and/or dusk are crepuscular
The tilted Earth orbits the sun every 365.25 days
All activity patterns/rhythms are innate and are controlled by an internal clock. The clock is set by an environmental cue known as a zeitgeber. The clock is synchronised by cues to determine the time of onset (and offset) of activity.
When the organism's activity is set to a particular zeitgeber, the the activity is said to be entrained.
Needs to be continuously entrained
Activity rhythms under the control of an internal clock are known as endogenous and these rhythms display a characteristic period.
In the absence of environmental factors (ie, when conditions are constant) the clock still runs and the organism still displays rhythmic activity. The period of the activity in constant conditions is known as the free-running period.
Circadian
Circannual
Circatidal
Circalunar
Circasemilunar
A few activity patterns are determined by external factors only and are known as exogenous rhythms (photosynthesis)
When constant conditions cease through the introduction of a zeitgeber, the organism's activity starts to phase-shift - the start of the activity moves each day until it concides with the onset of the zeitgeber.
The internal clock is under genetic control, with several genes identified in a range of organisms.
In humans, daylight is detected by the eyes and the message transmitted to the SCN in the hypothalamus of the brain, which is the site of the clock. This may then stimulate or inhibit the production of the hormone melatonin by the pineal gland (daylight inhibits production of melatonin, promoting wakefulness). In humans, the free-running period of the internal clock is about 24.5 hours, and therefore it constantly needs to be reset to entrain to the 24-hour day using the light/dark cycle as a zeitgeber.
The adaptive advantage of having activity controlled by environmental cues entraining an internal clock is that it allows an organism to anticipate or predict the onset of favourable conditions, therefore gaining maximum advantage of these.
Synchronisation may occur between species
Flowering
Flowering in many angiosperm flowers is an annual event and occurs in response to changing photoperiod.
Plants are placed into one of the following three groups, depending on the photoperiod that induces flowering.
Short-day plants - need a period of long nights (a photoperiod of less than a certain critical day length, CDL) before they can flower. Therefore these plants lower in autumn, winter or early spring.
Long-day plants - need a period of short nights (a photoperiod of greater than a CDL) before they can flower. Therefore these plants flower in late spring into summer.
Day-neutral plants - flower independently of the length of the photoperiod. Therefore these plants flower all year / any time of the year (typical of plants that live in the tropics).
This ability of plants to respond to the photoperiod is controlled by a light-sensitive pigment called phytochrome, which is present in the leaves and exists in two different forms, R(red)/P(r) (inactive form) and P(far-red)/P(fr) (active form), which can change from one to the other. P(red) absorbs red light and is converted to P(far-red), while P(far-red) absorbs far-red light and is converted to P(red). As sunlight contains more red light than far-red light, during the day P(red) converts to P(far-red), while during the night the unstable p(far-red) slowly reverts back to P(red).
If the day is long enough, P(far-red) accumulates and long-day plants flower.
If the night is long enough, P(red) accumulates and short-day plants flower.
Coincides with activity of animal pollinators and the long, warm days provide extended hours for pollinator activity - therefore chances of pollination are maximised.
The long photoperiod and warm temperatures increase photosynthesis and therefore promote growth of buds and flowers, therefore increasing flowering success.
Increased flower production and pollination opportunities increase chances of successful reproduction of the species.
If the start of the activity is getting later each day, the free-running period is slightly greater than 24 hours.
If the start of the activity is getting earlier each day, the free-running period is slightly less than 24 hours
k strategy
Courtship behaviour
Pair bond
A territory is a defined area containing a nest site set up by a group defending against all others of their own species
Provides a comparatively safe place for mating, nesting and rearing offspring
Increases the chances of successfully reproducing, rearing young and surviving.
Territory costs
Energy demanding for monitoring and defending the boundaries
Potential physical risks/injuries resulting from boundary disputes
A home range surrounds a territory. It is shared with other groups of the same species and is where they forage for food. The home range is not defended.
Reduced the risk of predation, as there are fewer night-active predators.
Work together in maintaining territory.
Solo parenting
Offspring is left on its own for a period of time, so there is a greater risk of death through heat loss, and they are more vulnerable to predation.
Reduces energy demands of each individual.
Mutualism is a relationship between members of two species in which both benefit (+ and +), therefore the survival chances of the individuals in the relationship increase (compared with the survival chances of organisms if they are not in the relationship).
Exploitation is a relationship between members of two species in which one benefits (+) while the other is harmed (-).
Parasitism is a relationship in which one individual (the parasite) feeds off another living individual (the host). The parasite benefits (+), while the host is harmed (-).
Parasitism is a very common and successful lifestyle and parasites exist in most animal groups (parasitism is rare in plants).
Types Of Parasites
Ectoparasites are free-living and feed on the host from the outside, typically piercing the skin and feeding by sucking the host's blood.
Endoparasites live inside the host, typically in the blood system feeding on blood or in the digestive tract feeding on digested food.
Parasites display specific adaptations for their lifestyle.
Endoparasites typically have a complex life cycle involving two different hosts (carnivore and herbivore). This allows for distribution of the species / reduction in infraspecific competition / reduced chances of over infestation of the host (which may kill it - parasitoidism)
Social (or brood) parasitism is known to occur when individuals of a species use the services of another species to raise their young.
Predation is a relationship in which one individual (the predator) hunts, kills and consumes and individual of another species (the prey). The predator benefits (+), while the prey is harmed (-).
Predators display specific adaptations for hunting/killing, while the prey displays counteradaption to reduce the risk of predation (such reciprocal adaptations are an example of co-evolution.
Camouflage
Cryptic colouration
Eye positioning
Speed/strength/stamina
Poison
Teamwork
Warning colouration occurs in a variety of animals and indicates to predators that they are unpalatable, harmful, toxic, or venomous
Herbivory is a relationship in which on individual (the herbivore) feeds on plant material. The herbivore benefits (+), which the plant is harmed (-).
Herbivores are not predators.
Herbivores display specific adaptations to their lifestyle, while plant often display adaptations to reduce/deter herbivory such as thorns/spines and toxic chemicals.
Interspecific Relationships
Mutualism
Commensalism
Ammensalism
Interspecific Competition
Interspecific Competition occurs when resources become limited. Members of both competing species are harmed (- and -), with reduced survival chances and reduced reproductive success.
The more similar the resource requirements of the two species are (ie, the greater the niche overlap), the more intense the competition is.
The more limited the resource(s) are, the more intense the competition is.
May also act to limit the extent of another species' niche, and if the niche requirements are two similar, then one species may outcompete and eliminate the other (Gause's Principle).
Plants may exploit herbivores in order to get pollinated. Pollinators are attracted to the plant by colour/chemicals/shape to obtain nectar from the flower. In doing so, the herbivore accidentally collected pollen on its body, which is transferred to the next flower/plant - pollination occurs.
Exploitation - Allelopathy
In this response, cells in the roots secrete a chemical (an allelochemical) into the soil that inhibits the growth of surrounding plants.
Reduces interspecific competition from these plants for resources.
The plant that releases the chemicals benefits (+), while the inhibited plants are harmed (-).
An example of antibiosis.
Groups are typically organised to a degree and are co-operative in that the individuals assist one another in various ways.
Breeding
Rearing of young
Defence against predators
Access to food
Forming and maintaining a group depends on behavioural adaptations of the individuals. These behaviours have been selected for, as they enhance survival and reproductive success.
The advantages of group living must outweigh the disadvantages if the group is to continue.
Advantages
Better defence against predators; young put on inside of herds.
Greater ability to find food, especially as a result of co-operative hunting
Mates readily available, increasing breeding success
Assistance with rearing young.
Greater learning opportunities
Population control by controlled access to mates/food
Protection from physical environment
Greater ability to spot predators, as more eyes/ears are on lookout.
Disadvantages
Increased intraspecfic competition, especially as group gets larger - this can result in poor health / reduced reproductive success / increased mortality.
Increased risk of the spread of disease resulting from close contact with others in the group.
Controlled access to mates, so some individuals do not get to breed (especially males).
Intraspecific competition
Occurs when resources become limited. As individuals of the same species have the same resource requirements, infraspecific competition is typically more severe than interspecific competition.
Competition for resources may provoke aggression between individuals - this is commonly seen in competition between males for mates during the breeding season.
Agonistic behaviour
Agonistic behaviour in groups results in the reduction of aggression.
This is seen in distinct postures/gestures/vocalisations that threaten another animal. These behaviours are understood and the threatened animal typically backs off / backs down or displays submissive behaviours, with the result that serious aggressive encounters are rare.
Agonistic behaviours tend to be ritualised, with dominant and submissive behaviours recognised and acted on by other members of the group.
Hierarchies occur when individuals have a specific rank (or status) within a group.
Ranking determines an individual's access to resources such as food, water and mates, therefore acts to reduce intraspecific competition, as well as aggression.
Only high-ranked individuals get to breed. Therefore only the alleles of the stronger individuals enter the gene pool / occur at higher frequency in the gene pool.
Ranking is determined by any, or a combination, of:
Sex
Age
Experience
Size
Strength
Rankings are typically determined by trials of strength. They may change over time.
The boundaries are defined and may be marked.
Vocalisations and/or visual displays may be used to advertise the location of a territory, warn off others of the species and/or attract a mate.
The size and location of a territory are determined by the area that the owner(s) can successfully defend, as well as the strength and/or status of the owners(s).
Males unable to secure a territory typically do not secure a mate and breed. Therefore, their alleles do not enter the gene pool. Territories therefore act as population controls and enhance the gene pool, as only the strongest individuals get to breed and contribute their successful alleles to the gene pool.
r strategy
Increases the chances of working together successfully to mate and rear offspring.
Large numbers of eggs are produced with little investment of time and energy in raising the offspring.
This strategy provides little risk to the parents, but has a low success rate.
Few offspring are produced with a large investment of time and energy in raising the offspring.
This strategy creates a higher risk to the parents, but is offset by a higher success rate.
A pair bond forms between the male and female as a result of courtship behaviour.
Courtship behaviour ensures that mating occurs between individuals of the same species and so maintains the isolation of the gene pool.
Courtship behaviours tend to be a series of repeated, ritualised activities and responses, typically initiated by the male.
Courtship also allows for sexual selection of males by females. The males display their size/strength/health/beauty and the female selects the 'best' - ie, the male that has the 'best genes', increasing the chances of her offspring having similar favourable traits.
In some species, a lek system exists in which males congregate in an area and display to watching females.
Typically only a few males will get to breed with the females, so contributing their favourable alleles to the gene pool.
Several different mating systems occur.
Monogamy
Polygyny
Polyandry
Polygynandry
Co-operative breeders
In which numerous male/female partnerings occur within the social group and most adult assist with raising the offspring.
Adults do not typically raise the offspring.
Synchronised spawning
In which males and females release gametes simultaneously, often in response to environmental cues, and fertilisation is external.
Both parents can share the duties of raising/feeding/protecting the young, which reduces the demands (eg, energy) on each parent and increases the chances of successfully raising the young until they are independent.
The main disadvantage is that the male does not have access to a range of potential partners, and therefore in unable to contribute his alleles in higher frequency to the gene pool.
Parental care refers to the behaviour associated with caring for and raising for young.
The advantage of providing parental care is that the survival chances of the offspring until they are independent are improved.
The disadvantages of parental care is the costs to the parents - energy demands are high when feeding and protecting the young. In addition, the parents are at increased risk of harm/death when they protect the young from potential predation and/or weather events.
Allogrooming benefits the group as it establishes/reinforces social bonds and reduces the risk of aggression.
Benefits young when they hatch by providing them with a warm protective environment surrounded by food, therefore increasing their survival chances.