Structure of woody plants
Vascular cambium
Initiation of the vascular cambium
Ray initials
Fusiform initials
Arrangement of cambium cells
Secondary xylem
Growth rings
Heart Shaped & sapwood
Reaction wood
Secondary phloem
Outter bark
Cork & the cork camium
Lenticels & oxygen diffusion
Initiation of cork cambia
Secondary growth in roots
Anomalous secondary growth
Unusual primary growth
Vascular cambium is the main growth tissue in the stems and roots of many plants, specifically in dicots such as buttercups and oak trees, gymnosperms such as pine trees, as well as in certain vascular plants. It produces secondary xylem inwards, towards the pith, and secondary phloem outwards, towards the bark.
are isodiametric cells about equal in all dimensions and they produce the vascular rays, which constitute the horizontal system of secondary tissues; this horizontal system acts in the translocation and storage of food and water.
are elongate cells that produce the conducting cells in both the secondary xylem and secondary phloem and the other cells in the axial system.
Ray & fusiform initials are organized in specific patterns. Ray initials are grouped together in short vertical rows and only one cell wide. Fusiform initials occur in regular horizontal rows
Storied cambium
Nonstored cambium
the initials are arranged in horizontal regular file having a stratified structure. The initials are usually short and all of about the same length. The ends of the initials occur approximately at the same level
the fusiform initials and rays are not arranged in horizontal tiers on tangenti.
Periclinal wall
cell is one that is parallel to the surface of the plant body. A periclinal division is one that results in the formation of periclinal walls between daughter cells. Such a division results in an increase in girth of the organ
Vascular cambium
Fascicular cambium
Interfascicular cambium
the main growth tissue in the stems and roots of many plants, specifically in dicots such as buttercups and oak trees, gymnosperms such as pine trees, as well as in certain vascular plants. It produces secondary xylem inwards, towards the pith, and secondary phloem outwards, towards the bark.
cambium present between the xylem and phloem of a vascular bundle
The portion of the vascular cambium that develops from parenchyma cells
All cells formed to the interio of the vascular cambium develop into the secondary xylem
Annual ring in wood, some are produced 2 in 1 year and some as late as 9 years
Wood that is formed in response to mechanical stress
a center of a log is always darker in color than the outer wood, and drier and more fragrant. The darker wood is heartwood and lighter, moister outer region is sapwood
type of phloem that forms from the vascular cambium during the secondary growth. The secondary growth is responsible for the growth in girth in plants, especially trees
Cork cambium differs greatly from vascular cambium in both structure and morphogenic activity. All of its cells are cuboidal, like ray initials. after each division the inner cell almost always remains cork cambium whereas the outer cell differentiates into a cork cell
The outter most dead layers of bark, from the surface to the innermost cork cambium.
Cork/Phellem cell
non-living and has cell walls that are composed of a waxy substance that is highly impermeable to gases and water called suberin. The layer of dead cells formed by the cork cambium provides the internal cells of the plants with extra insulation and protection.
Periderm
the corky outer layer of a plant stem formed in secondary thickening or as a response to injury or infection.
Phelloderm
) A part of the periderm made up of cells produced inwardly by the cork cambium
The impermeability of cork has a negative as well as advantageous consequences. It keeps out pathogens and retains water it also blocks absorption of oxygen interfering with the respiration of sapwoods, vascular cambium and inner bark.
The first bark on young stems differs from bark formed when the stem is older, if the first cork cambium arises by reactivation of the epidermal cells the first outer bark contains only periderm and cuticle that is very smooth
Any form of secondary growth that does not conform to that typically occurring in gymnosperms and eudicots
Included phloem
In certain types of anomalous secondary growth, patches of secondary phloem may be located within the secondary xylem
Such as in palms, seedlings initially grow wider while remaining short, during this establishment growth the seedling establishes additional adventitious roots