Community Ecology
Diversity
Predator-Prey Interactions
Beneficial Interactions
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communities range from a few species to thousands
a community is a group of species that occur together at the same place and time #
these species are limited by each other through competition, predation, or changes they produce in the environment as well as disturbances from accidents or humans
beaver ponds are a community that experience rapid change in plant type and growth, and animal activity
as beavers and other new organisms mold the pond habitat it experiences stages of succession - predictable sequence of change
spruce-fir forests endure for thousands of years, they are an example of climax communities - stable self-maintaining communities
the effects humans have on communities has been devastating, #
community restoration projects are necessary to reverse some of these effects
human activities are constantly resulting in habitat loss and fragmentation
a checklist, or species count can measure the richness of a community
ecology usually focuses on several organisms rather than all of them because there are too many to quantify
the objectives of ecology studies can vary, one might cover species checklists, while another cover just the herbs shrubs and trees present
studies could examine tropic levels, or even alleles present in a community
scale is important to consider in ecology
local - small area less than a few square kilometers
region - area larger than a few square kilometers but typically much less than a continent, encompasses all members of many species, and speciation and extinction are important
biome - a large region characterized by the dominant plants present and climate. Ex. temperate grassland, temperate rainforest, tropical rainforest, desert, and salt marcsh
biogeographical region - an extremely large region that usually coincides with a continent or other very large region bounded by a geological or climatic barrier to migration barriers such as oceans and mountains prevent migration of most species #
species area relationship
whitaker proposed 3 scales: alpha for local, beta for differences between several small sites, and gamma is the number of species within a region
a checklist and species are relationships dont take abundance of each species into account, a species abundance distribution would plot the amounts of each species
diversity goes down with increase in lattitude, this may be due to harsher conditions and less rainfall or shorter growing seasons farther north
50 million years ago, most of the earth's climate was tropical. 20 million years ago the earth started to cool and ice sheets formed. so temperate plants began to evolve, this probably relates to tropic diversity - temperate plants have had less time to delineate
ex. of difference in species abundance between two biogeographical regions
diversity and abundance bring richness to communities
One predator, one prey
in nature this doesnt happen, but this model works for understanding harvesting fish, lumber or deer #
most populations follow a logistical growth curve
as a plant species reaches its carry capacity the herbivore and plant will cycle up and down together
the populations of both predator and prey can fall together or even both become extinct if all of the prey is consumed
the functional response includes a predator's feeding rate and the handling time to consume the prey
this functional response is prey dependent, it is faster when prey are more dense
Lotka-volterra single predator prey model
population stability is represented by dN/dT or zero growth isocline
Rosenzweig-MacArthur includes more factors than lotka-volterra
more complex formulas, and a curved zero growth isocline
sometimes enriched numbers of prey result in predators overconsuming prey and losing food source. this is called the paradox of enrichment
we can choose to consume just enough of a prey to keep populations stable at a point called the maximum sustained yield
fixed effort harvesting, setting a limit on time and style of harvest, like a 2 month long hunting season, where bag limit can vary based on population size
fixed quota harvesting - limit is a specific number no matter how long it takes.
in japan and norway they are harvesting whales even though they are endangered.
predator selection with multiple prey
most communities have multiple plant species that are prey to herbivores
optimal foraging theory studies why herbivores choose certain plants over others
optimal diet model
first predators evolve to prefer whichever prey yields the most energy per unit of handling time
second if high yield prey become scarce the predator will have more success with a broader diet and include prey that are less efficient but more abundant
third, some items like grasses will always be eaten even though lower in nutrients and other never eaten because of defense like cactus spines
fourth the probability a plant will be eaten is dependent on the abundance and efficiency of other plants
this is more representative of nature
interspecies competetion
several species compete for the same resources
in exploitation competition the resource is consumed making ti less available for other organisms
in interference competition, one organism restricts another's access to resources even though the first might not be using it
invasive species have the ability to increase their population density despite competition
a resource is any substance or factor that can lead to growth
bracken fern is an intense competitor, throwing shade on shrubs below it yet able to grow in shade if something manages to grow taller than it
if all herb populations are low and one spikes, the increase in prey from its boom could result in more consumption of the lower populated species as well, endangering their survival, this is apparent competition
mutualism is when two organisms interact and both are benefitted
ex of mutualism is pollinators and plants they polinate
facilitation is when one organism helps another without benefit to themselves
these ants feed on the nectar and are protected by spines, they also keep out other insects and their wastes and decomposition enrich the soil
solid mathematical models for mutualism dont exist
there are small costs for mutualism, something must be given up on each side
facilitation ex is nurse plants, who make a small habitat for other plants underneath
facilitators serve a role in succession
metapopulations
all of the populations of a species are separated physically and geographically
if local populations are connected by migration and gene flow the local pops are a metapopulation
metapopulation model assumptions
a region of the environment is composed of many discrete patches in which the species can live
some patches are occupied by the species, whereas others are not
empty patches will become colonized by migration from occupied patches
populations within individual patches have a probability of going extinct within that patch
high quality patches are source habitats and low quality ones a sink habitat
as long as their are new patches an ephemeral species can survive, but if limited to currently occupied patches they may die out
empty patches are not surplus or unneeded, some species need to rotate through them if they are adapted to early succession and poor in later
a fugitive species is one that needs to colonize a new patch before it dies out an occupied one
Food Chains and Webs
communites have at least 3 trophic levels
primary producers (plants, algae, cyanobacteria)
primary consumers (herbivores)
and secondary consumers (carnivores)
possibly tertiary consumers if plant richness occurs
this direct line of consumption is called a food chain
if we trace out all predators and prey we will create a network of interactions called a food web
energy flow web will maps out energy consumptions and is difficult for real communities
food chain and web diagrams are common but problematic and lack variables
keystone species are those that have a dramatic effect on a community
in the northern pacific ocean the sea otter was almost lost due to hunting #
this keystone species was necessary to keep sea urchins in check
without sea urchin predation, urchins over grazed kelp preventing it from thriving
humans have reduced alot of predators that consumed herbivores, allowing plant life to thrive
a common thread in this chapter is the negative impact humans have on natural habitats
this is a common thread throughout the chapter, we must study ecology because we have such a negative impact on it