Community Ecology

Diversity

Predator-Prey Interactions

Beneficial Interactions

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communities range from a few species to thousands

a community is a group of species that occur together at the same place and time #

these species are limited by each other through competition, predation, or changes they produce in the environment as well as disturbances from accidents or humans

beaver ponds are a community that experience rapid change in plant type and growth, and animal activity

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as beavers and other new organisms mold the pond habitat it experiences stages of succession - predictable sequence of change

spruce-fir forests endure for thousands of years, they are an example of climax communities - stable self-maintaining communities

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the effects humans have on communities has been devastating, #

community restoration projects are necessary to reverse some of these effects

human activities are constantly resulting in habitat loss and fragmentation

a checklist, or species count can measure the richness of a community

ecology usually focuses on several organisms rather than all of them because there are too many to quantify

the objectives of ecology studies can vary, one might cover species checklists, while another cover just the herbs shrubs and trees present

studies could examine tropic levels, or even alleles present in a community

scale is important to consider in ecology

local - small area less than a few square kilometers

region - area larger than a few square kilometers but typically much less than a continent, encompasses all members of many species, and speciation and extinction are important

biome - a large region characterized by the dominant plants present and climate. Ex. temperate grassland, temperate rainforest, tropical rainforest, desert, and salt marcsh

biogeographical region - an extremely large region that usually coincides with a continent or other very large region bounded by a geological or climatic barrier to migration barriers such as oceans and mountains prevent migration of most species #

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species area relationship image

whitaker proposed 3 scales: alpha for local, beta for differences between several small sites, and gamma is the number of species within a region

a checklist and species are relationships dont take abundance of each species into account, a species abundance distribution would plot the amounts of each species

diversity goes down with increase in lattitude, this may be due to harsher conditions and less rainfall or shorter growing seasons farther north

50 million years ago, most of the earth's climate was tropical. 20 million years ago the earth started to cool and ice sheets formed. so temperate plants began to evolve, this probably relates to tropic diversity - temperate plants have had less time to delineate

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ex. of difference in species abundance between two biogeographical regions

diversity and abundance bring richness to communities

One predator, one prey

in nature this doesnt happen, but this model works for understanding harvesting fish, lumber or deer #

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most populations follow a logistical growth curve

as a plant species reaches its carry capacity the herbivore and plant will cycle up and down together

the populations of both predator and prey can fall together or even both become extinct if all of the prey is consumed

the functional response includes a predator's feeding rate and the handling time to consume the prey

this functional response is prey dependent, it is faster when prey are more dense

Lotka-volterra single predator prey model

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population stability is represented by dN/dT or zero growth isocline

Rosenzweig-MacArthur includes more factors than lotka-volterra

more complex formulas, and a curved zero growth isocline

sometimes enriched numbers of prey result in predators overconsuming prey and losing food source. this is called the paradox of enrichment

we can choose to consume just enough of a prey to keep populations stable at a point called the maximum sustained yield

fixed effort harvesting, setting a limit on time and style of harvest, like a 2 month long hunting season, where bag limit can vary based on population size

fixed quota harvesting - limit is a specific number no matter how long it takes.

in japan and norway they are harvesting whales even though they are endangered.

predator selection with multiple prey

most communities have multiple plant species that are prey to herbivores

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optimal foraging theory studies why herbivores choose certain plants over others

optimal diet model

first predators evolve to prefer whichever prey yields the most energy per unit of handling time

second if high yield prey become scarce the predator will have more success with a broader diet and include prey that are less efficient but more abundant

third, some items like grasses will always be eaten even though lower in nutrients and other never eaten because of defense like cactus spines

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fourth the probability a plant will be eaten is dependent on the abundance and efficiency of other plants

this is more representative of nature

interspecies competetion

several species compete for the same resources

in exploitation competition the resource is consumed making ti less available for other organisms

in interference competition, one organism restricts another's access to resources even though the first might not be using it

invasive species have the ability to increase their population density despite competition

a resource is any substance or factor that can lead to growth

bracken fern is an intense competitor, throwing shade on shrubs below it yet able to grow in shade if something manages to grow taller than it

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if all herb populations are low and one spikes, the increase in prey from its boom could result in more consumption of the lower populated species as well, endangering their survival, this is apparent competition

mutualism is when two organisms interact and both are benefitted

ex of mutualism is pollinators and plants they polinate

facilitation is when one organism helps another without benefit to themselves

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these ants feed on the nectar and are protected by spines, they also keep out other insects and their wastes and decomposition enrich the soil

solid mathematical models for mutualism dont exist

there are small costs for mutualism, something must be given up on each side

facilitation ex is nurse plants, who make a small habitat for other plants underneath

facilitators serve a role in succession

metapopulations

all of the populations of a species are separated physically and geographically

if local populations are connected by migration and gene flow the local pops are a metapopulation

metapopulation model assumptions

a region of the environment is composed of many discrete patches in which the species can live

some patches are occupied by the species, whereas others are not

empty patches will become colonized by migration from occupied patches

populations within individual patches have a probability of going extinct within that patch

high quality patches are source habitats and low quality ones a sink habitat

as long as their are new patches an ephemeral species can survive, but if limited to currently occupied patches they may die out

empty patches are not surplus or unneeded, some species need to rotate through them if they are adapted to early succession and poor in later

a fugitive species is one that needs to colonize a new patch before it dies out an occupied one

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Food Chains and Webs

communites have at least 3 trophic levels

primary producers (plants, algae, cyanobacteria)

primary consumers (herbivores)

and secondary consumers (carnivores)

possibly tertiary consumers if plant richness occurs

this direct line of consumption is called a food chain

if we trace out all predators and prey we will create a network of interactions called a food web

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energy flow web will maps out energy consumptions and is difficult for real communities

food chain and web diagrams are common but problematic and lack variables

keystone species are those that have a dramatic effect on a community

in the northern pacific ocean the sea otter was almost lost due to hunting #

this keystone species was necessary to keep sea urchins in check

without sea urchin predation, urchins over grazed kelp preventing it from thriving

humans have reduced alot of predators that consumed herbivores, allowing plant life to thrive

a common thread in this chapter is the negative impact humans have on natural habitats

this is a common thread throughout the chapter, we must study ecology because we have such a negative impact on it

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