x=age, Nx=number of individuals alive at age x, Sx=proportion of individuals of age x that that survive to age x+1, Ix=proportion of individuals that survive from birth (0) to age x, Fx= average number of offspring born to a female while she is age x, IxFx= average number of offspring per capita at age x, Ix(Fx)(x)= average number of offspring per capita at time x, weighted by age x