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Course Diagram, Ch 22 Seed Plants I: Seed Plants Without Flowers ("…
Course Diagram
Ch 22 Seed Plants I: Seed Plants Without Flowers ("Gymnosperms")
Division Gnetophyta
similar leaves to dicots
mostly vines or small shrubs
Division Ginkgophyta: Maidenhair Tree
ancestors are not known
leaves turn yellow in autumn
single living species
Division Cycadeoidophyta: Cycadeoids
almost identical to cycads
had vegetative features
ALL EXTINCT
Division Cycadophyta: Cycads
100 species
9-10 genera
do not ovules
frequently confused with ferns
Division Coniferophyta: Conifers
proembryo
other cells develop into embryo
suspensor
pushes other cells deep into megagametophyte
ovuliferous scale
auxiliary bud and megasporophylls fused laterally
cone bracts
short axis bears leaves
compound cones
shoot with auxiliary buds
simple cones
single short unbranched axis that bears microsporophylls
two different types of shoots
short shoot
long needle leaves
long shoot
tiny papery leaves occur here
550 species
50 genera
Division Pteridospermophyta: Seed Ferns
Three Divisions
Cycadeoidophyta
cycadeoids, all extinct
Cycadophyta
cycads, extant
Pteridospermophyta
seed ferns all extinct
earliest seed fern appeared in Upper Devonian Period
Division Progymnospermophyta: Progymnosperms
progymnosperms
later gave rise to conifers, cycads, and other gymnosperms
Evolution of Seeds
pollen chamber
holding area
micropyle
hole in integument that permitted sperm cells to swim to the egg
integument
layer of tissue that projects upward
difficult but not impossible
Archaeopteridales
reproduction was heterosporous
more derived progymnosperm
Aneurophytales
little webbing between branches
vary from shrubs to large trees
relictual progymnosperms
Ch 8 Structure of Woody Plants
Anomalous Forms of Growth
Unusual Primary Growth
establishment growth
form of primary growth
palm trees
Anomalous Secondary Growth
sweet potatoes
storage roots
alternative cambia produce secondary bodies that differ from common type
Secondary Growth in Roots
storage capacity of a woody root can be increased
perennial roots also form bark
root vascular cambium has ray and fusiform initials
Outer Bark
Initiation of Cork Cambia
timing is more variable
Lenticels and Oxygen Diffusion
lenticels
regions of aerenchymatous
Cork and the Cork Cambium
periderm
layers of cork cells
phelloderm
cell matures into layer of parenchyma
cork cell
outer cell
cork cambium/phellogen
new cambium
Secondary Phloem
axial system
responsible for conduction up and down stem or root
has axial and radial system
formed from vascular cambium
Secondary Xylem
Reaction Wood
response to stress
Heartwood and Sapwood
sapwood
lighter, moister outer region
heartwood
dark wood
Growth Rings
diffuse porous
vessels located throughout it
ring pores
vessels restricted to early wood
annual ring/ growth ring
1 year growth
early wood
first wood formed
Types of Wood Cells
two basic types of ray parenchyma
procumbent cells
no direct connection with axial cells
up right cells
softwoods
few or no fibers
hardwood
wood of all basal angiosperms and eudicots
radial system
develops from ray initials
axial system
tracheary elements
longitudinal conduction of water through wood
derived from the fusiform
secondary xylem=wood
all cells formed interior to vascular cambium
Vascular Cambium
Arrangement of Cambial Cells
nonstoried cambium
without any horizontal pattern
storied cambium
evolved more recently
regular horizontal rows
Ray Initials
produce storage parenchyma
similar to fusiform
short and cuboidal
Fusiform Initials
perpendicular cell division
increase number of cambia cells
anticlinal walls
parallel cell division
periclinal wall
long taper cells
Initiation of the Vascular Cambium
only has two types of cells
ray initials
fusiform initials
interfascicular cambium
connects on each side with fascicular cambia
fascicular cambium
continue dividing
fascicle= bundle
woody species
one of the meristems that produce secondary plant body
Ch 16 Genetics
Mutations
Inversion
put in backwards
Insertion
addition of extra DNA
Deletion
pieces of DNA is lost
Point Mutation
single base is converted to another base
smallest mutation affecting least amount of DNA
Mutation
any change in DNA
DNA Repair Process
repairing can cause serious problems
must be neither too efficient nor too ineffective
Somatic Mutations
eventually becomes sexual cell
happen in cells that never lead to sex cells
Effects of Mutations
natural selection eliminates the deleterious one and preserves the beneficial one
mutations are almost always harmful
depends on
extent
position
nature
Causes of Mutations
transposon
carries genes that code for proteins
like insertion sequence
Insertion Sequences
& splicing it into DNA somewhere else
contains genes that code for enzymes
involved in cutting insertion sequence
Mutagen
something that causes a mutation
Replication of DNA
ligated
attached with covalent bonds
replication fork
DNA coils and separates looks like fork :fork_and_knife:
semiconservative replication
each DNA strand acts as template for making the complementary strand
DNA Polymerase
primer RNA act as DNA-synthesizing enzyme
primer RNA
ribonucleotides are polymerized into short pieces
replicon
small bubble where strands separate from each other
DNA replicated in S phase
Monohybrid Crosses
Cross
sexual reproduction between individuals
Multiple Alleles
multiple triple alleles
subscripts are uses
polymorphic
900 existing sites
Test Cross
must be made on annual plans
pure-bred lines
homozygous dominant
Test Cross
cross involving one cross known to be homozygous recessive for the trait being considered
complete dominance
presence of one allele completely makes presence of homologous allele
Monohybrid Crosses with Complete Dominance
recessive
other possible phenotype of the heterozygote that is less likely to be than the dominant
dominant
phenotype of the heterozygote is like the parent of the effective alleles
Crossing Heterozygous with Themselves
Punnett Square
does not represent outcome of any one cross, just posibilility
test that can be used to test genotypes and phenotypes
Selfing
plant's own pollen is used to fertilized its eggs
Monohybrid Crosses with Incomplete Dominance
incomplete dominance
heterozygous phenotype differs from both homozygous phenotypes
heterozygous
two different alleles for one gene
homozygous
two identical alleles for a gene
F2
offspring of F1
F1
offspring of the parents
parental generation
parents
monohybrid cross
single character is analyzed and studied
Dihybrid Crosses
cross in which two genes are studied and analyzed simultaneously
Genes on the Same Chromosome: Linkage
map unit
one equals 1% of probability that crossing over will occur between them
measure of separation of genes on a chromosome
recombinant chromosomes
formed from crossing over of the homologous chromosomes and recombination of alleles
parental type chromosomes
linkage group
set of genes that do not undergo independent assortment, being part of one chromosome
linked genes
genes located close to each other on a chromosome undergo crossing over only rarely, so they're linked
Crossing Over
Genes on Separate Chromosomes: Independent Assortment
independent assortment
alleles of one gene moved independently of the alleles of the other genes
Other Aspects of Inheritance
Multiple Sets of Chromosomes and Gene Families
gene family
various copies happen through duplication and may evolve independently
all located within a single haploid genome
set of several copies of an ancestral gene
paralogs
genes within single species that evolved from same ancestral gene
polypoid
plants with more than two sets of chromosomes
Lethal Alleles
its presence can kill a plant
can be difficult to detect in early stages
Maternal Inheritance
variegation
pattern of spots, stripes, or patches in leaves or other organs, caused by plastid mutations
pollen parent and ovule parent :question:
uniparental inheritance
zygote obtains all its plastids and mitochondrion genomes from maternal parent
(Maternal Inheritance)
biparental inheritance
alleles of both parents are transmitted equally to progeny
Multiple Genes for One Character: Quantitative Trait Loci
pleiotropic effects
multiple phenotype effects of one mutation
quantitative trait loci
phenotypes that vary in degree and can be attributed to polygenic effects
epistasis
having multiple genes for each trait
Ch 9 Flowers and Reproduction
Fruit Types and Seed Dispersal
Classification of Fruit Types
indehiscent
fruit does not break open and release seed
dehiscent fruit
break open and release seeds
fleshy fruit
fruits that are eaten during the natural seed distribution process
dry fruit
not typically eaten by natural seed distributing animals
True Fruits and Accessory Fruits
multiple fruit
all individual fruits of a florescence fuse into one fruit
aggregate fruit
separate carpels of one gynoecium fuse during development
simple fruit
fruit develops from a single ovary
accessory fruit
used if any nonovarian tissue is present
false fruit
true fruit
refers to fruits containing only ovarian tissue
Inflorescences and Pollination
inflorescence
two basic arrangements occur
indeterminate inflorescence
determinate inflorescence
give collective visual signals to pollinators
discrete group of flowers
Flower Structure and Cross Pollination
Ovary Position
half inferior
sepals, petals, and stamen are intermediate
superior ovary
ovary located above sepals, petals, and stamen
inferior ovary
ovary located below sepals, petals, stamen
Wind-Pollinated Flowers
Animal-Pollinated Flowers
coevolution
zygomorphic
bilaterally symmetrical flower
actinmorphic
synonym for regular flowers
results in highly specific interaction
type of evolution in which two species become increasingly adapted to each other
Monoecious and Dioecious Species
dioecy
condition in which a species has two types of sporophyte
monoecy
condition of having staminate flowers located on the same plants as carpellate flowers
nonessential organs
sepals and petals do not produce spores
perfect flowers
has both essential organs
imperfect flowers
flowers that lack either or both essential organs
essential organs
produce critically important spores
Stigma and Pollen Incompatibility
compatibility barriers
chemical reactions between pollen and carpels that prevent pollen growth
Cross-Pollination
self pollination
or another flower of a same plant
pollination of a carpel by pollen from the same flower
is the pollination of a carpel by pollen from different individual
Sexual Reproduction
Fruit Development
pericarp
entire fruit wall
Three layers become distinct during growth
endocarp
innermost layer
mesocarp
middle or flesh
exocarp
outer layer, skin or peel
fruit
what ovary matures to
Embryo and Seed Development
exalbuminous
endosperm is sparse or absent at maturity
albuminous seed
mature seed in which endosperm is abundant
hypocotyl
root/shoot junction
epicotyl
embryonic stem
radicle
embryonic root
cotyledon
leaflike structure involved in either nutrient storage
suspensor
pushes the embryo deep into the endosperm
Fertilization
endosperm
nourishes the development of the zygote
double fertilization
sperm fusion with egg nucleus and the other with polar nuclei
karyogamy
fusion of the nuclei
plasmogamy
fusion of protoplasts of the gametes
Gametophytes
embryo sac
technical term for multinucleate megagametophyte
pollen tube
pollen grain germinates with tube-like process that carries sperm to egg vicinity
generative cell
cells that give rise to sperm cells
vegetative cell
cells that do not give rise to sperm cells
Flower Structure
carpels
have three main parts
ovary
where megaspores are produced
style
elevates stigma to useful position
stigma
catches pollen grains
constitute the gynoecium
located at the highest level on receptacle
pollen
microspores and microgametophytes
stamens
two parts
anther
where pollen is produced
filament
its stalk
collectively called androecium
above petals
perianth
sepals and petals
petals
corolla
all petals together
above sepals
sepals
calyx
all sepals together
lowermost and outermost of four floral appendages
incomplete flowers
lack one of four main types
complete flowers
have all four types
sepals, petals, stamen, and carpels
receptacle
other flower parts are attached
pedicel
flower stalk
The Plant Life Cycle
alternation of generations
sporophyte or gametophyte
life cycle of plants
macrogametes
large eggs
do not swim
microgametes
sperms cells
swim
gametophyte
haploid plant that produces gametes
spores
where meiosis occurs
sporophytes
have organs
always diploid
gametes
egg
female
sperm
male
diploid adults have sex organs that produce haploid sex cells
Asexual Reproduction
fragmentation
large spreading or vining plant grows to several meters length and individual parts become self-sufficient by establishing adventitious roots
Ch. 11 Respiration
Types of Respiration
Photorespiration
wastes 30%of energy from photosynthesis
occurs when RuBP carboxylase adds oxygen
Anaerobic Respiration
Produces ATP & NADH
respiration without oxygen
lactic acid build up
glycolysis
breaks down glucose
Respiration of Lipids
fatty acids broken down
by acetyl CoA
or enters citric acid cycle
synthesis of carbs
Aerobic Respiration
Citric Acid Cycle/Krebs Cycle
Benefit
MORE ATP :smiley:
respiration with oxygen
respiration with oxygen
glycolysis
occurs in:
plastids
cytosol
produces:
NAD+
NADH
ATP
first in anaerobic too :open_mouth:
Pentose Phosphate Pathway
occurs in:
cytosol and plastids
synthetic pathway is more important than respiratory energy
important source for fundamental compunds
Heating-Generating Respiration
also called Thermogenic Respiration
studied through cyanide
poisons aerobic respiration
body temperature
NADH converts to heat
plants produce better than human
Fermentation of Alcoholic Beverages
Warnings
ethanol
can be used as a cleaner
harms
DUI = 5th greatest cause of death
death
benefits
lowers heart disease
fermentation produces ethyl alcohol
is a depressant :frowning_face:
Spirits
alcohol 20% ethanol
distilled
doable because different liquids boil at diff temp
alcohol content ^^^
solution= toxic to yeast
Wine
15,000 varieties
6000 BCE
fermented grapes
continues as long as sugar is present
20-30% weight = sugar
Beer
How to:
68oC-73oC stand for hours
moisten grains and germinate
fermenting starchy cereal grains
rice
sake= fermentated
barely
greatest # of enzymes
= faster than others
MOST COMMON
corn
wheat
Respiratory Quotient
RQ rise
photosynthesis meeting anabolic needs
extra respiration
in dark
RQ= low
helps measure amounts of gases exchanged
respiratory metabolism
indicator of the type of substrate being respired
Total Energy Yield of Respiration
36 ATP TOTAL
Aerobic respiration
2 NADH
glycolysis yields 2 ATP
Anaerobic respiration
NET ATP =2
2ATP start process
4 ATP are synthesized
Environmental and Internal Factors
Internal Regulation
different cells have different respiration rate
seeds with dormant period
no respiration
mature fruit
respiration bursts
fruit maturation
respiration remains steady
mature embryo
respiration decrease
Temperature
above 40oC respiration decreases
Above 30oCincreases
Below 50oC respiration decreases
increase 10oC doubles respiration rate
Lack of Oxygen
ANOXIA
no oxygen :forbidden:
HYPOXIA
Lesser amounts of oxygen
plants
1-2% maintain full respiration
Ch 12 Transport Processes
Long-Distance Transport: Xylem
Control of Water Transport by Guard Cells
light and CO2are normal controlling factors
if moisture content is adequate
primarily powered by water loss in atmosphere
Water Transport Through Xylem
embolism
expands until surface encounters a solid barrier
sometimes called air bubble
space between two portions
cavitation
hydrogen bonding broken over large region and water column breaks
poikilohydry
body water content that changes with habitat moisture
transcuticular transpiration
water loss through cuticle
transstomatal
water loss through stomata
cohesion-tension hypothesis
molecules cohere sufficiently to withstand tension
water is pulled upward by transpiration
most widely accepted
Properties of Water
cohesive
behaves weakly when bound together
frozen
molecules become strongly bound together
acts on neighboring cells
adhesive
water molecules interact with many other substances
Long-Distance Transport: Phloem
callose
seals certain regions
growing pollen tubes
damaged sieve elements
P-protein
fine network adjacent to plasma membrane
phloem
sinks
sites that receive transported phloem sap
specific mass transfer
mass transfer/cross sectional area of phloem
mass transfer
sugars and nutrients transported by phloem by the hour
STM/CC complex
in phloem loading and conduction
sieve tube members and guard cells work together
polymer trap mechanism
conducting cell plasma membranes are permeable to monosaccharides and disaccharides but not polysaccharides
sources
sites from which water and nutrients are stored
pressure flow hypothesis
flow in phloem is due to active loading in sources and active unloading in sinks
Short-Distance Intercellular Transport
apoplast
volume of plant not occupied by protoplast
intercellular spaces and cell walls of plant
symplast
protoplasm of one plant, one continuous mass
Transfer Cells
found where rapid, short distance transport occurs
sugar is load in or out of phloem
passing nutrients to embryos
glands that secrete salt
ridge-like growths on inner surface
walls are smooth on outer surface
Motor Cells
similar to guard cells
accumulate or expend K+
cells at joints
Guard Cells
when open= high negative osmotic potential
abundant K+
& adjacent cells are in equilibrium when stomata are fully opened and closed
water enters and leaves cells at same rate
cat eye when open
at night = shrunken
opening and closing of stomatal pores
Water Potential
Water Movement
Water Potentials must always be considered in pairs or groups
If water potentials are at equilibrium there is no movement of water
Water moves whenever there is a difference in water potential within the mass of water
matric potential
decrease water's free energy
water's adhesion to nondisolved structures
osmotic potential
number of particles present in solution
effect that solutes have on water potential
pressure potential
measure in megapascals (MPa)
can be positive
effect pressure has on water potential
water potential
increased by:
elevated
under pressure
heat
free energy of water
Cells and Water Movement
plasmolyzed
cell constantly losses water
incipient plasmolysis
protoplast has lost just enough water to pull away from wall
lysis
Plant cell NEVER bursts :red_cross: :forbidden:
animal cells explodes in places in pure water
Diffusion, Osmosis, and Active Transport
Intracellular transport
vesicles migrate through the cytoplasm and fuse with another organelle
molecular pumps
use ATP to forces molecules across membrane
ACTIVE TRANSPORT
aquaporins
membrane protein that permits water to cross membrane rapidly
3 Membrane Types
Selectively Permeable
allow certain substances to pass through
Completely Permeable
membranes do not allow anything to pass through as occur as isolation barriers
Freely Permeable
membranes allow all solutes to diffuse through them and have little biological significance
osmosis
diffusion through a membrane
diffusion
random movement of particles in solution causes them to move from areas where they are in lower concentration
Ch 20 Nonvascular Plants: Mosses, Liverworts, and Hornworts
Division Anthocerotophyta: Hornworts
The Gametophyte Generation
many different shapes
have numerous chambers
distinctive
Division Hepatophyta: Liverworts
The Sporophyte Generation
elaters
single, elongated cells with spring shaped-walls
The Gametophyte Generation
archegoniophores
producing eggs
antheridiophore
umbrella shaped outgrowth
air pores
not stomatas
thallus
a body without roots, stems, and leaves
divided into two basic groups
thallose liverworts
leafy liverworts
resembles moss
Division Bryophyta: Mosses
Metabolism and Ecology
can improve the microhabitat for seedlings
important in later establishment
two critical factors
The Sporophyte Generation
calyptra
layer of cells derived from the neck of the archegonium
peristome teeth
one or two sets of teeth like structure around the mouth of sporangium
seta
between foot and sporangium, narrow stalk
operculum
lid like top of a sporangium
capsule
upper cell grows by cell division and expands into simple apical sporangium
foot :footprints:
zygote of moss and basal cell
The Gametophyte Generation
archegonia
structure that produces egg
antheridia
produces sperm cells in ascomycete fungi
protonema
grow from spore during germination
leptoids
resemble sieve cells
hydroids
innermost cortex
rhizoids
penetrate surface of substrate
gametophores
form dense mounds
tightly appressed
grow tightly together
leafy stems
mosses
grow from apical meristem
thrive in many places within cities
perennial
Classification of Nonvascular Plants
hornworts-Anthocerotophyta
mosses-Bryophyta
Liverworts-Hepatophyta
not known how closely related moss, liverworts, and hornworts are
Characteristics of Nonvascular Plants
can grow to be very large
embryophyes that do not have vascular tissue
people are very familiar with
Ch 23 Seed Plants II: Angiosperms
Eudicots
Asterid Clade
two small orders
campanulids
lamiids
most derived
Rosid Clade
two large clades
malvids
fabids
named for the rose order rosales
Basal Eudicots
perisperm
surrounds developing embryo
nutritive tissue
betalains
water soluble pigments
Monocots
liliales
Alismatales
Tepal
members of perianth when it is not certain if they are really sepals or petals
Commelinoid Monocots
have unique types of epicuticular wax
differ from the others in several unusual synapomorphies
Basal Angiosperms
uniaperturate
ancestral condition for angiosperms
pollen that only has a single germination pore
contain living descendants of several groups that originated while angiosperms were still a small clade
Classification of Flowering Plants
basal angiosperm
not newly discovered
several clades of angiosperms that arose before the rest of the angiosperms diverged into monocots and eudicots
almost all angiosperms are classified as monocots or eudicots
eudicot
clade of angiosperms that contains most species formally known as dicots
monocot
informal term for any member of the flowering plant class Liliopsida
Changing Concepts About Early Angiosperms
paleobotanists and taxonomists think the transition from gymnosperm to angiosperm happened in the Jurassic Period
most botanists said that angiosperms were monophyletic
generalized flower
it has all parts
sepals, petals, stamens, and carpels
ranalean flower
magnolia type flower was thought to be relictual
hypothesis developed by C. E. Bessey
Ch 26 Community Ecology
Interconnectedness of Species: Food Chains and Food Webs
energy flow web
difficult to make for real communities
keystone species
species that dramatically effects the structure of its community
number of individuals
portion size
food web
various organisms eat or decompose others, and these in turn are consumed
food chain
direct line of consumption
Metapopulations in Patchy Environments
assisted dispersal
animals are capture in one area and released into another
fugitive species
species that survives by colonizing new patches, flourishing temporarily, then colonizing more patches and dies out in old one
sink habitat
low quality
source habitat
high quality patch
metapopulation
makes four assumptions
populations within individuals patches have a probability of going extinct within that patch
empty patches will become colonized by migration from occupied patches
some patches are occupied by the species whereas other suitable patches are not
a region of the environment is composed of many discrete patches in which the species can live
population that consists of several local populations interconnected by migration and gene flow between patches
Beneficial Interactions Between Species
primary succession
organism becomes established on newly created substrates
nurse plants
spiny desert shrubs
presence of some plants creates conditions that allow other plants to become established
facilitation
one organism helps another without receiving any benefit
first organism facilitates
mutualism
two organisms interact with that both organisms benefit
Predator-Prey Interactions
Apparent Competition
predator may prey on several species
Competition Between Species
resource
any substance or factor that can lead to increased growth rate as its availability is increased and that is consumed by an organism
interference competition
one organism restricts another organism's access to resources even though the first might not using it.
exploitation competition
occurs when organism actually consume a shared resource making it less available for organism
Predator Selection Among Multiple Prey
optimal diet model
makes four predictions
optimal foraging theory
examine the interactions between these factors in an attempt to understand why herbivores eat the plants they do while ignoring the others
One Predator, One Prey
fixed quota harvesting
limits damage to a species by setting limits on amount that can be harvested
fixed effort harvesting
technique that limits damage to a species by automatically decreasing harvests
maximum sustained yield
amount of organisms population that can be harvested without population decreasing
paradox of enrichment
loss of a population from a habitat as a result of that habitat being altered to benefit the population
zero growth isocline
line indicating population stability
Lotka-Volterra model
criticized for being to simplistic
dP/dt=faNP-qP
rate of change of predator number
dN/dt=rN-aNP
rate of change with time of prey population
prey-dependent
predator's functional response depends on prey density
functional response
predator's feeding rate and handling time
Diversity
Diversity and Latitude
could change in temperatures were the same long enough
equator
benign environment
higher latitude has more severe conditions
far north
lower diversity than closer to the equator
different plants depending on environment on earth
Diversity and Scale
species abundance distribution
plot of the number of species in a region into abundant classes
measuring diversity
gamma
number of species within a region
beta
compares differences between smaller sites in large region
alpha
number of species or growth forms that occur at local site
Level of Scale
Biogeographical region
Biome
Region
Local
species-area relationship
S=cA^z
relationship between area and species richness
larger communities more diverse than small
Ch 5 Tissues and the Primary Growth of Stems
Stem Growth and Differentiation
Primary Growth
growth and formation from apical meristem
Primary Tissues
tissues produced by apical meristems
provascular tissues
young cells of xylem and phloem
Protoderm
epidermal cells in early stages for differentiation
protophloem
exterior cells
metaxylem
largest tracheary elements
suapical meristem
cells dividing and growing
producing cells for region below
below apical meristems
apical meristems
stems grow longer by creating new cells at their tips
Internal Organizations of Stems: Arrangement of Primary Tissues
Vascular Bundles
primary phloem
all are collateral
xylem and phloem strands running parallel
pith
region of parenchyma similar to cortex
interior to cortex
primary xylem
primary plant body
Phloem
two types of conducting cells
"sieve element"
must be alive to conduct
parenchyma cells
sieve tubes members
companion cells
sieve cells
albuminous cells
Xylem
conducts water and minerals
scalariform thickening
provides more strength
annular thickenings
secondary wall organized as rings
two types of conducting cells
vessel elements
tracheids
tracheary elements
Vascular Tissues
not circulatory system
phloem
distribute sugars and minerals
xylem
conduct water and minerals
Cortex
cells are compact
photosynthetic parenchyma
interior to epidermis
Epidermis
trichomes
hairs
Cutin
can inhibit CO2 from entering
Stroma fixes this
fatty substance that makes cell impermeable to water
shields internal cell
other leaves or stems
dust
animals
barrier against invasion
single layer of living parenchyma
Outermost surface of stem
External Organizations of Stems
Rhizomes
Tubers
storing nutrients
potatoes
grow only short period
fleshy horizontal stems
allow plant to spread underground
phyllotaxy
important to not shade each other
arrangement of leaves on stem
terminal bud
extreme tip of each stem
Leaf Axil
Axillary Bud
young leaves
atypical meristem
miniature shoot
stem area just above point where leaf attaches
Nodes
Internodes
regions between nodes
where leaves are attached
Shoot
stem + leaves, flower, &/or bud
Stem
Axis
Basic Types of Cells and Tissues
Sclerenchyma
Two Types
Conducting
vascular cell
transports water
Mechanical
sclereids
cuboidal
fibers
strong
supports plant itself
does not droop
detrimental
snow
animals
wind
elastic
goes back to original shape
both
thick secondary cell wall
primary cell wall
Collenchymara
produced on shoot tips and young petioles
require a lot of glucose
long and flexible
grape stems :grapes:
Relatable to clay
can be deformed by pressure & retains new shape :open_mouth:
plasticity
mainly thin
except in corners
Parenchyma
easy to build
involves little glucose
Some die at maturity
SUBTYPES
Transfer
increase surface area through extensive knobs
short distance transport
Chlorenchyma
numerous chloroplasts
thin walls
involved in photosynthesis
Grandular
transports:
minerals
sugar
few chloroplasts
secret:
oils
resins
mucilage :question:
fragrances
nectar
makes up
seeds
fruits
petals
soft leaves
most common type of cell and tissue in plants :!!:
remain thin
only have primary wall
Ch 6 Leaves
External Structure of Foliage Leaves
parallel venation
veins run side by side
veins
bundles of vascular tissue
compound leaf
blade divided into several individual parts
simple leaf
one part blade
sessile leaf
long and narrow
petiole
stalk
holds blade into light
leaf blade= lamina
ventral surface
smooth
adaxial side
dorsal surface
abaxial side
large veins protrude like back bones
blade's lower side
flat light-harvesting
nutritious
obvious function = photosynthesis :sunflower:
Morphology and Anatomy of Other Leaf Types
Insect Traps
entire leaf blade wraps around insect
passive traps
incapable of movement
active traps
move during capture
trap and digest insects
Leaves with Kranz Anatomy
CO 2 transport
C4 photosynthesis
Tendrils
cells are capable of sensing contact with an object
grow indefinitely
modified leaf
Spines
protection
modified leaves of auxiliary buds
Bud Scales
produce thin layer of corky bark
primary role is protection
rarely compund
small
form tight layer around stem tip
bud scales
most common modifications
Leaves of Conifers
perennial
simple
sclerophylls
THICK
hypodermis
epidermis
cuticle
Sclerophyllous Foliage Leaves
sclerophylls
perennial
leaves could live for two years
leaves are soft, flexible, and edible
must produce more sugars by photosynthesis
could lose energy every time it made a leaf
Succulent Leaves
photosynthesis occurs deep
mesophyll has little air spaces
mesophyll transparent
water loss through stomata
leaves are cylindrical or spherical
reduced CO2 intake
optimal surface to volume shape
desert habitat
Internal Structure of Foliage Leaves
Petiole
stipules
small and die early
contribute to photosynthesis
protect shoot apical meristem
two small flaps
different bundles patterns
leaf traces
one, three, five or more vascular bundles
transition between stem and lamina
varies in size
Vascular Tissues
bundle sheaths
many fibers become arranged
eudicot
midrib= midvein
between palisade parenchyma and spongy mesophyll
Mesophyll
spongy mesophyll
lower portion of leaf
palisade parenchyma
main photosynthetic tissue of most plants
upper surface of most leaves
ground tissues interior to the leaf epidermis
Epidermis
leaves
hairy
protection
must be reasonably water proof
transpiration
water loss through epidermis
Initiation and Development of Leaves
Monocots
constant basal expansion
lamina becomes broad and expanded
can be linear and strapped
apical meristem
adjacent to primordium
Basal Angiosperm and Eudicots
leaf primordium
extends upward, narrow cone
bulk of mid rib
increases in thickness
cells just interior to protoderm growing outward
Ch 7 Roots
Other Types of Roots and Root Modification
Roots of Strangler Figs
can kill host
penetrate rapidly
grow various angles
possibly doesn't touch soil for years
nutrients from rainwater
birds eat and deposit onto other trees
Haustorial Roots of Parasitic Flowering Plants
haustoria
grows around small branch
not all related
roots of parasitic plants
Root Nodules and Nitrogen Fixation
infection thread
penetrates one cell after another to the root cortex
nitrogen fixation
chemical conversion of nitrogen into usable compounds
Mycorrhizae
endomyocorrhizal association
cannot pass casparian strip
penetrate root cortex
herbaceous
ectomycorrhizal relationship
fungi penetrate outermost cortex but do not invade cell themselves
woody forest plants
Contractile Roots
root contraction = anchored shoot in soil
contraction caused by change in cortex cells
uppermost portion slowly contract
more contraction that prop roots
Aerial Roots of Orchids
velamen
water proof barrier
several layers of large dead cells
white
rain forest
on surface on bark and dangle freely in air
epiphytic
living attached to the branches of trees
Prop Roots
buttress roots
brace trunk against the wind
upper side grows more rapidly
place tension on stem and act as stabilizers
transport additional nutrients and water to stem
bases of corn and grasses
Storage Roots
long term storage for carbs
used to produce new shoot
Internal Structure of Roots
Mature Portions of the Root
root pressure
powerful water absorption and water pressure
passage cells
once thought to be passageways for mineral absorption
root hairs function for just days and die
Zone of Maturation/ Root Hair Zone
pericycle
lateral roots
vascular tissue and parenchyma cells form irregular region
casparian strips
control types of minerals that enter xylem water stream
bands of altered walls
endodermis
cylinder cells
greatly increasing absorption of water and minerals
root hairs grow outward
Zone of Elongation
inner cell is provascular tissue
outer most cell is protoderm and epidermis
tissues are permeable
cells expand greatly
behind root apical meristem
Root Apical Meristem
quiescent center
becomes active is meristem is damaged
form new apical meristem
mitotically inactive central region
central cells not synthesizing DNA
orderly
Root Cap
constantly regenerating itself
cells pushed towards edge of cap
small cells
meristematic
specific structure
Origin and Development of Lateral
destroys cells of cortex
busts through cell
initiated by cell division in pericycle
External Structure of Roots
Structure of Individual of Roots
root hairs
increase root's surface area
form only on part of root that is not elongating
root hair zone
lateral roots emerge
epidermal cells extend out as trichomes
zone of elongation
cells undergo division and expansion
mucigel
lubricates passage of root through soil
complex polysaccharide
secreted by root cap cells
root cap
constantly worn away
protects root apical meristem
Organization of Root Systems
adventitious roots
increase absorptive and transport capacities
fibrous root system
similar sized roots
radicle
usually becomes largest root
embryonic root that develops from taproot
lateral roots/branch roots
small
enormous absorptive surface
Ch 17 Population Genetics and Evolution
Evolution and the Origin of Life
The Presence of Life
Oxygen
created conditions that selected for the evolution of aerobic respiration
allowed earth to rust
Early Metabolism
energy metabolism
glycolysis
Aggregation and Organization
are not postulated to have been alive
inheritable
formed at random
Formation of Polymers
Chemicals Produced Chemosynthetically
meteorites
various organic compounds from the moon
Conditions on Earth Before the Origin of Life
energy sources
second atmosphere
reducing atmosphere
produced by release of gases from the rock matrix composing Earth
Speciation
Convergent Evolution
ex=cacti and euphorbias
convergent evolution
evolution of two phenotypically distinct species, organs, that they strongly resemble each other
Divergent Speciation
genetic drift
in a small gene pool, the alteration in allele frequencies mostly by accidents rather than natural selection
adaptive radiation
species rapidly diverges into many new species over a short time
hybrid sterility
two populations occasionally interbreed or are artificially cross-pollinated and produce viable seed
postzygotic internal isolation barriers
same as prezygotic but occurs AFTER
prezygotic isolation mechanisms
phenomena that prevent successful interbreeding of two populations but act so early that fertilization is not possible
sympatric speciation
when two groups become reproductively isolated even though they grow together
biological reproductive barriers
any biological phenomenon that prevents successful gene flow
allopatric
if speciation occurs
abiological reproductive barriers
any physical, nonliving feature that prevents two population from exchanging genes
Phyletic Speciation
gene flow
vegetative propagation
seed dispersal
pollen transfer
the movement of alleles physically through a space
Rates of Evolution
difficult to identify the presence of particular alleles in a population
very few mutations cause new phenotypes
most populations are relatively well adapted to their habitat
Population Genetics
deals with the abundance of different alleles within a population and the manner in which the abundance of a particular allele increases, decrease, or stays the same
Multiple Selection Pressures
need for pollinators
cold
drought
insect attack
Situations in which Natural Selection Does Not Operate
sides of a road are cut
competition does not occur
cannot happen if all individuals are identical
Factors that Cause the Gene Pool to Change
natural selection
2 ways for it to occur
progeny must differ from each other in their types of alleles
population must produce more offspring that can possibly grow and survive to maturity in that habitat
the most significant factor causing the gene pool changes is usually described as survival of the fittest
artificial selection
artificial mutation
crop plants and domestic animals
process in which humans purposefully change the allele frequency of a gene pool
Accidents
ex. large meteorite hits earth
an organism cannot adapt
Mutation
existing decrease while new increase
occurs constantly
Ch 21 Vascular Plants Without Seeds
The Term "Vascular Cryptograms"
vascular cryptograms
lack seeds so reproduction is hidden
indicates they have vascular tissue
ferns and fern allies
The Megaphyll Line of Evolution: Euphyllophytes
Ferns
leptosporangia
forms a small outward protrusion
initiated when a single surface cell divides periclinally
eusporangium
initiated when surface cells undergo periclinal divisions
sori
clusters of sporangia where meiosis occurs
leaf trace
vascular bundle that extends from the stem vascular bundles through the cortex and enters a leaf
leptosporangiate ferns
more than 12,000 species
familiar to us
leaf gap
area above leaf trace where there is no conducting tissue
Equisetophytes
monopodial growth
main trunk, lateral branches, true leaves, true roots
sporangiophore
umbrella shape
Monilophytes
lignophytes
woody plants
two sister clades
Origin of Megaphylls (Euphylls)
euphyllophytes
megaphyllous plants form a monophyletic clade
sporophyll
sporangium-bearing, leaf-like structure
webbing
concept that lamina originated by the production of parenchyma cells between the telomes
planation
occurs in one plane, resulting in flat system
telome
twigs, last of dichotomy
telome theory
theory that leaves evolved from branching systems by overtopping, planation, and webbing
homoplastic structures
megaphylls
leaves that evolved from branch systems
enations of zosterophyllsophytes
leaves on gametophytes
Trimerophytes
pseudomonopodial branching
single main trunk rather than a series of dichotomies
overtopping
trimerophytes had unequal branching and one stem was more vigorous
The Microphyll Line of Evolution: Lycophytes
Extant Genera
ligule
small flaps of tissue
Heterospory
cones/strobili
sporangia clustered together in compact groups
Morphology
microphylls
present in lycophytes
type of leaf that evolved from an enation
Early Vascular Plants
Zosterophyllophytes
enations
thought to have been the ancestor of leaves in lycophytes
small projecting flap of tissue
another group of early vascular plants
Rhyniophytes
siphonostele
later evolved, pith in center
exarch protostele
protoxylem on edges of several groups next to phloem
metaxylem located in center of xylem
endarch protostele
metaxylem differentiates on outer edge of xylem mass
protoxylem is located in center
protostele
solid mass of xylem and no pith
rhyniophytes
earliest vascular plants
equal dichotomous branching
both branches of equal size and vigor
cooksonia
earliest fossils that were vascular plants
Ch 10 Photosynthesis
Environmental and Internal Factors
Crassulacean Acid Metabolism
appears in
euphorbias
lilies
bromeliads
orchids
cactus family
selectively advantageous
hot dry climate
not particularly efficient
occurs at night
metabolic adaptation improves conservation of water
C4 Metabolism
advantageous in warm dry conditions
25oC-30oC
happens in Kranz anatomy
monocots in hot climates
mechanism CO2 is absorbed and O is kept away
photorespiration
energy wasting process
Water
CO2 is gone water cant react
especially at night
stomata close
very important to photosynthesis
Leaf Structure
depends on where plant is located
less stomata
also lengthen photosynthesis
some have water loss reductions
can lengthen photosythesis
Light
Duration
summer=better :sunflower:
Number of hours a day sunlight is available
Quantity
also consider shape and size of plant
intensity or brightness
greatest at noon
Quality
colors or wavelength
Photosynthesis
Anabolic Metabolism
gluconeogenesis
anabolic synthesis of glucose
storage compunds
store starch=
chloroplasts and amyloplasts
long term storage
starches
most important
intermediate term storage
sugars
short term storage
ATP NADPH
Stroma Reactions
operate 3-phosphoglyceraldehyde and take out of chloroplasts
builds anything the plant needs
nucleic acids
amino acids
fats
sugars
(Calvin/Benson Cycle)
ATP and NADPH interacts with CO2=carbs
The Light Dependent Reactions
Photosystems 2
P680
Photosystems 1
P700
absorb red light
Electromagnetic Radiation Spectrum
accessory pigments
most common
carotenoids
chlorophyll b
molecules that strongly absorbs wavelength not absorbed by chlorophyll a
VISIBLE LIGHT
pigment
CRITICAL
chlorophyll a
absorbs quanta
absorbs certain wavelength and has specific color
400-750 nm
radiowaves
microwaves
infrared lights
uv light
x-rays
gamma rays
Water and light create intermediate ATP and NADPH
Energy and Reducing Power
Other Electron Carriers
Plastocyanin
chloroplast membrane
holds copper
small protein
Plastoquinones
dissolve into chloroplasts membrane
hydrophobic
pick up 2 e, bind 2 p
Cytochromes
carries electrons short distance
in chloroplasts thylakoid
holds iron "heme"
small proteins
Reducing Power
reducing power=ability to force electrons onto compounds
Earth atm = 21% oxygen
electron lost=oxidized
oxidizing agents= take electrons away
oxygen
increases positive charger
electrons added=reduced
reducing agents=give electrons away
hydrogen
reduces positive charger
oxidative state=number of e- during redox reactions
covalent bonds
oxidized=atoms do not carry as many electrons as it could.
Energy Carriers
3 methods ADP>ATP
oxidative phosphorylation
occurs in all of plant
ADP phosphorylated to ATP
substrate-level phosphorylation
high energy phosphate groups
photophosphorylation
occurs only in chloroplasts
light energy in photosynthesis
energy enters world through photosynthesis
Ch 25 Populations and Ecosystems
The Structure of Ecosystems
Tropic Levels
energy flow
decomposers
break down remains of other organisms
fungi and bacteria
secondary consumers
carnivores
primary consumers
herbivores
primary producers
autotrophs
basically feeding levels
Species Composition
refers to number and diversity of species that coexist in an ecosystem
Temporal Structure
changes that an ecosystem undergoes with time constitute
Physiognomic Structure
physical size and shape, and their distribution in relation to each other and physical environment
The Structure of Populations
r- and K-Selection
K-Selection
douglas firs, bristle cones
r-selected species
annuals or small shrubby
Age Distribution: Demography
carrying capacity
number of individuals in each population that can live in a particular ecosystem is limited
symbolized by K
biotic potential
number of offspring produced by an individual that live long enough to reproduce under ideal conditions
generation time
affect population growth
length of time from the birth of one individual until the birth of its first offspring
age distributions
=demography
relative young portions, middle aged, and old individuals
Geographic Distribution
Local
uniform distributions
all individuals are evenly spaced
occur in orchards and tree plantations
clumped distributions
spacing between plants is either large or small but rarely average
random distribution
no obvious identifiable pattern to the position of individuals
Boundaries
Limiting factor
various nutrients
light
water
Plants in Relationship to Their Habitats
Operational Habitat
aspects of the habitat that definitely affect a plant
Habitat
set of conditions in which an organism completes its life cycle
Biotic Components of the Habitat
Organisms Other than Plants
grazing
eats herbs
browsing
eating twigs and plants
predation
relationship in which one species benefits and the other is harmed
commensal relationships
one species benefits and the other is unaffected are also common between plants and animals
Other Plant Species
transplant experiment
plants are transplanted to another spot to live in common ground with another plant
ecotypes
race of a species that are adapted to a particular environmental factors in certain parts of the species range
niche
the set of conditions exploited best by one species
competitive exclusion
species that is less adapted is excluded from the ecosystem by superior competitors
competition
disadvantageous for both organisms
mutualism
interaction is beneficial for both organisms
Plant Itself
Abiotic Components of the Habitat
Disturbance
Latitude and Altitude
Soil Factors
C horizon
parent rock and rock fragment
B horizon
zone of deposition
A horizon
zone of leaching
uppermost
pioneers
must tolerate several conditions
first plant that invade a new soil
Climate
tolerance range
between the low and high extremes