Population Genetics and Evolution

concepts

population genetics

rates of evolution #

speciation

evolution & life origin

factors that change

gene pool

4 factors

artificial selection

natural selection

accidents

mutation

some situations

natural selection

does not operate

multiple selection pressures

occurs two ways

phyletic speciation

requires

movement of alleles

3 ways

seed dispersal

vegetative propagation

pollen transfer

divergent speciation

reproductive isolation

causes

abiological reproductive barriers

biological reproductive barriers

special type

adaptive radiation

convergent evolution

earth conditions

before life

atmospheric chemicals

energy sources

time available

chemosynthetically produced chemicals

formation of polymers

aggregation and organization

early metabolism

oxygen

presence of life

evolution

gradual conversion of

one species to

one new species

several new species

occurs by

natural selection

slow process

may require

thousands of generations

millions of years

Earth's age #

billions of years

discovered

in

independently by

mid 1800s

Alfred Russel Wallace

Charles Darwin

"survival of fittest"

terms

population genetics

abundance of alleles

within a population

and manner

in which

a particular allele

increases

remains the same

decreases

gene pool

number of alleles

in all

gametes in population

ratio remains same

if only

sexual reproduction considered

demonstrated by

G. H. Hardy

G. Weinberg

occurs continually

allele frequencies

new alleles increase

existing alleles decrease

significance depends on

population size

events that organisms

cannot adapt to

examples

meteorite collision

eliminates

organisms and alleles

may change

allele frequencies

volcanic eruptions

infrequent

hail storms

droughts

floods

continental drift

avalance

grazing animals

highway construction

when humans

purposefully change

allele frequency

in gene pool

examples

selective breeding of

crop plants

domestic animals

select

favorable traits

examples

high protein content

ability to survive

resistance to disease

in seeds

in

less fertilizer

less water

most significant factor

for occurrence

2 necessary conditions

population must

produce more offspring

than can survive

in that habitat

progeny must

affected by

predators

pathogens

limited resources

competitors

differ from others

in

types of alleles

factors not involved

purpose

intention

planning

voluntary decision

examples

genetically identical population

if adaptation impossible

no competition

or universal survival

would occur

in plowed fields

cut road sides

recently

burned area

flooded area

often

more than one

factor

change with

environment

time

examples

simultaneously

insects

weather

fungi

efficiency of

metabolism

water absorption

pleiotropic effects

may occur

"side effects"

some advantageous

some disadvantageous

depends on habitat

depends on habitat

typically

gradual

slow

most populations are

already well adapted

to their habitat

are especially

slow in

rapid in

ferns

Hawaiian asters

now 3 genera

new species

every 500,000 years

if

disruptive mutation

can happen

quickly

constructive mutation

can happen

slowly

when natural selection

causes

new species

to evolve

labeled this if

no fertile offspring

when two organisms

are crossed

aka gene flow

pollen grains

carry haploid genome

travels

by

wind

long distances

animal-mediated pollination

examples

birds

insects

dispersal mechanisms

long distance

short distance

example

fruits fall

next to parent

carried by

wind

floods

stream flow

small mobile species

that reproduce

vegetatively

if gene flow

does not

keep species homogeneous

are

physical nonliving features

that prevent

gene exchange

between two populations

when two species

grow separately

if speciateion results

examples

mountains

deserts

oceans

UV light

dry air

are

any biological phenomena

that prevent

successful gene flow

examples

prezygotic isolation mechanisms

discrimination pollinators

affected by

differences in

a flower's

fragrance

shape

flowering timing

allopathic speciation

grow together

aka geographic speciation

sympatric isolation

evolutionary changes

in pollinators

color

occurs before

zygote can form

environmental diversity

postzygotic

internal isolation barriers

hybrid sterility

when

artificial cross-pollination

produce viable seeds

but seed grows

into sterile plant

occasional interbreeding

results in

hybrid inviability

early death in

zygote

embryo

species rapidly diverges

into new species

in only

few million years

occurs when

species enter

new habitat

with little/no

competition

environmental stress

examples

Hawaiian Islands

Galapagos Islands

first offspring

resemble

founder individuals

gene pool is

extremely limited

new alleles

build up rapidly

genetic drift

rapid chantes

in gene pool

due to

higher subjectivity

to accidents

can occur

on mainland

if sudden

environmental changes

delete dominant species

when

two distinct species

have similar phenotypes

favored by

natural selection

examples

euphorbias

cacti

evolved in

American deserts

evolved in

African deserts

convergence of phenotypes

image #

hypothesis

chemosynthesis

models life origin

using two processes

chemical

physical

proposed by

two scientists

A. Oparin

J.B.S. Haldane

in Russia

in England

initially

hydrogen

second atmosphere

several gasses

examples

produced by

release from

Earth's rock matrix

heavy meteorite bombardment

methane

ammonia

hydrogen sulfide

water

early on called

reducing atmosphere

due to

lack of

molecular oxygen

presence of

powerful reducing agents

most intense

from sun

UV radiation

gamma radiation

heat

from

coalescence of

gas and dust

radioactive decay

electricity

lightning storms

lightning around

volcanoes

unlimited

due to

lack of

molecular oxygen

1.1 billion years

from

Earth solidifying

to

arising of life

experimental tests

performed in

1953

consisted of

boiling water

reducing atmosphere

at bottom

at top

resulted in

formation of

complex organic compounds

from

monomers in

the early ocean

into masses

with

metabolism

example

fatty, hydrophobic material

not alive

aggregates would be

complete heterotrophs

when nutrient scarcity

needed enzyme

to synthesize

scarce molecules

from abundant molecules

evolution of

glycolysis

evolution of

two things

chlorophyll a

photosynthesis

resulted in

2 consequences

world can rust

created conditions that

selected for

aerobic respiration

liberation of

oxygen

2.8 billion

years ago

resulted in

oxidizing atmosphere

gradual transitions from

completely inorganic compounds

to

living bacteria

chemistry is

more complex

than nonliving objects

no unique properties

from nonliving objects

Cross link description: The beginning of the chapter emphasizes that Earth is billions of years old. This is further underscored by the discussion about oxygen becoming liberated 2.8 billion years ago, which was after the inital formation of Earth.

Cross link description: Natural selection is the primary factor in altering the gene pool. The example of cacti and euphorbias which grow in completely different continents clearly demonstrates how traits best suited for an environment are considered more "fit" and survive for multiple generations until its gene frequency is higher than the former.

Cross link description: Evolution is generally a very slow process. However, in adaptive radiation when a species is introduced to a new habitat with little to no stress or competition, new alleles can build up relatively quickly. This results in more rapid divergence of species.

several types possible

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