RS 6
Amount and Rate of Blood Flow
- Rate of renal blood flow under basal conditions, approximately 1200 ml/min
(400 ml/100 g tissue/min), - is very high compared to other tissues.
- Total renal blood flow is approximately 20% of resting cardiac output,
- while the two kidneys make <0.5% of total body weight.
- Range of blood flow to kidney is rather narrow.
- It allows only a 25% increase (300 ml) over the basal renal blood flow of 1200 ml/min.
- This reserve is much higher in other organs.
- Higher blood flow to kidneys is related to
- Blood flow to the kidneys is directly proportional to the pressure difference between the renal artery and renal vein,
- and is inversely proportional to the resistance of the renal vasculature.
- Because the afferent arteriole, the efferent arteriole and the intralobular artery are
the major resistance vessels in the kidneys, they determine renal vascular resistance.
- In face of blood pressure changes,
- During exercise
- sympathetic tone to renal vessels increases
- and shunts renal blood flow to the skeletal muscles.
- the renal blood flow shows
remarkable constancy due to autoregulation.
- its excretory function rather
than its metabolic requirement.
Regulation of Renal Blood Flow
- The regulatory mechanisms affect the renal blood flow (RBF) and glomerular filtration rate (GFR) by changing the arteriolar resistance.
- Regulatory mechanisms of renal blood flow include:
- Autoregulation,
- Hormonal regulation and
- Nervous regulation.
Autoregulation of Renal Blood Flow
- Renal blood flow (RBF) and thus the glomerular filtration rate (GFR) remain constant over a wide range of renal arterial pressures
(80–200 mmHg).
- This occurs due to an intrarenal mechanism known as autoregulation.
- Autoregulation of RBF is accomplished by changing renal vascular resistance.
- When arterial pressure changes
(between 80–200 mmHg),
a proportionate change occurs in the renal vascular resistance which maintains a constant RBF.
Mechanisms of Autoregulation
Two mechanisms are considered responsible for autoregulation of RBF and GFR:
- one mechanism that responds to changes in arterial pressure,
- and another that responds to changes in NaCl concentration of tubular fluid.
- Myogenic mechanism.
- It is related to an intrinsic property of vascular
smooth muscle: - the tendency to contract when it is stretched.
- Thus, when renal arterial pressure is raised, the afferent arterioles are stretched, which contract and increase the vascular resistance.
- The increased vascular resistance offsets the effect of increased arterial pressure and thereby maintains a constant RBF and GFR.
- Tubuloglomerular feedback mechanism.
- Tubuloglomerular feedback (TGF) mechanism is based on the NaCl concentration of tubular fluid.
- It involves a feedback loop which operates as:
- Changes in the GFR cause changes in NaCl concentration of fluid in the loop of Henle.
- Changes in the NaCl concentration are sensed by the macula densa cells and converted into a signal.
- The signal from the macula densa cells changes the vascular resistance in afferent arterioles.
- Signals obtained due to increased concentration of NaCl produce vasoconstriction; conversely signals obtained due to decreased NaCl
cause vasodilatation of afferent arterioles.
- The effector mechanism responsible for vasoconstriction and vasodilatation is not exactly known.
- Perhaps, adenosine triphosphate
(ATP), which selectively constricts the afferent arterioles and metabolites of arachidonic acid, may contribute to TGF mechanism.