A key force in this evolution may have been the earth’s many cycles of environmental change, including, over the last 2.5 million years, more than 20 glacial cycles. The most recent of these was the melting and retreat of vast ice sheets about 15,000 years ago. One can imagine that, with every cycle of Pleistocene glaciation, migrants retreated to more southern breeding localities and made shorter migrations while the ice ruled the north. Then, when the ice receded, the migrants moved north to take advantage of renewed breeding opportunities in the unoccupied habitat that emerged.