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Vascular Plants Without Seeds ("Vascular Cryptograms" ("…
Vascular Plants Without Seeds
Early Vascular Plants
Rhyniophytes
Cooksonia
Earliest Vascular Land Plants
Now Extinct
Short and Simple Upright Stems
No Leaves
Equal Dichotomous Branching
Homosporous
Rhiniophytes
Rhynia
and
Aglaophyton
Similar to
Cooksonia
Prostrate Rhizome
Upright Naked Stems
Terminal Sporangia
Cuticle, Stomata, and Guard Cells
Lyonophyton
and
Sciadophyton
Gametophytes
No Sporangia
Larger and Complex than Common Mosses, etc.
Possible Alternate Phase of
Rhynia
Horneophyton
Sporophytes with Naked Axes
Stomata and Terminal Sporangia
Xylem Structures
Two Types of Organization
Prostele
Solid Mass of Xylem
No Pith
Endarch Prostele
Center Protoxylem
Outeredge Metaxylem
Exarch Protostele
Centered Metaxylem
Outside Groups of Protoxylem
Siphonostele
Pith Present in Center
Occurs in Ferns and Seed Plants
Zosterophyllophytes
Small Herbs
No Secondary Growth
Lateral Sporangia
Sporangia Opened Transversely
Exarch Prostele
Grew as Small Bunches
Upper Portions had Cuticle, Epidermal Cells, and Stomata
Lower Portion Had None of These Things
Significant Morphological Changes Occurred Quickly
Lycophytes
Morphology
Drepanophycus
and
Barawanathia
Similar to Presumed Ancestors
Earliest Lycophytes
Larger Enations
Called Mycophylls
Increased Photosynthesis
Single Trace of Vascular Tissue
Evolution of True Roots
Vascular Cambium and Secondary Growth
Wood Looked Like Secondary Xylem of Pines
Vascular Cambium Cells Couldn't Have Radial Longitudinal Division
Heterospory
Cones or Strobili
Clustered Sporangia in Compact Groups
Heterosporous
Having Both Microspores and Megaspores
Necessary Precondition for Seed Evolution
Shows Convergent Evolution
Extant Genera
All Are Small Herbs
Around 200 Species
Prostrate Rhizomes
True Roots
Short, Upright Branches
No Secondary Growth
All Are Homosporous
Lycopodium
Fairly Common
Also Called Ground Pine or Club Moss
Selaginella
Less common
Smaller Plants
Heterosporous
Ligule
Small Flap of Tissue
Isoetes
60 Species
Contains the Quillworts
Small, Corn-like Stem
Heterosporous
Euphyllophytes
Trimerophytes
Overtopping
Unequal Branching
Resemble Rhyniophytes
Pseudomonopodial Branching
Single Main Trunk
Small Lateral Branches
Branches Became More Uniform
Increased Vigor and Robustness
Origin of Megaphylls
Leaves that Evolved From Branch Systems
Present in All Seed Plants, Ferns, and Equisetophytes
Telome Theory
Megaphyll Evolution
Telomes
Ultimate Twigs
Last Dichotomy
Sporophyll
Sporangium-Bearing, Leaf-Like Structure
Euphyllophytes
Monophyletic Clade
Monilophytes
Two Sister Clades
Ferns and Fern Allies
Equisetophytes
Division Arthrophyta
Horsetails or Scouring Rushes
15 Extant Species
All Herbs
No Secondary Growth
Small Leaves
True Roots
Branches Sometimes Present
Distinctive Internal Structure
Stems have a Pith
Siphonsteles
Vessels are Rare
Contains Canals
Stomatal Pores and Guard Cells
Sporangiophore
Umbrella-Shaped
Short Stalk
Flat, Shield-Shaped Head
Compact Spirals Forming a Strobilus
Ferns
First Appeared in Devonian Period
12,000 Living Species
Living Species Called Leptosporangiate Ferns
Found in Almost Any Habitat
Leaves
Leathery or Delicate
Small or Large
Almost Always Compound
Distinct Apical Cell
Considerable Amount of Vascular Tissue
Dichotomous Veins
Photosynthesis and Sporophylls
Sori
Located Underside of Leaf
Clusters of Sporangia
Where Meiosis Occurs
Most Homosporous
Two Sporangia Types
Eusporangia
Several Surface Cells Undergo Periclinal Divisions
Fundamental Type of Sporangia
Outer Cells Develop into Sporangium Wall
Inner Cells Proliferate into Sporogenous Tissue
Relatively Large Sporangium, Many Pores
Nearly All Plants
Leptosporangia
Initiated by Single Surface Cell Dividing by Periclinal Division
Forms Small, Outward Protrusion
Only Few Spores Produced
Only Occurs in Leptosporangiatae
Small Set of Sporogenous Cells
Thing Covering of Sterile Cells
"Vascular Cryptograms"
"Ferns and Fern Allies"
Indicates Vascular Tissue
Reproduction is Hidden
Do Not Group them Formally
Refers to a Grade
"Byrophyte"